Only Jeremy Chardy progressed of the five seeded players in action in Montpellier on Wednesday, with Tsonga’s 6-2 7-6 (7-3) defeat of fellow Frenchman Gilles Simon the most notable result.
The former world number five will now face another compatriot in the quarter-finals as he continues to rebuild his career, with Chardy up next after a straight-sets win over qualifier Antoine Hoang.
David Goffin was involved in the late match of the day, but the second seed was brushed aside by Filip Krajinovic 6-4 6-4.
Benoit Paire was also beaten by Tomas Berdych, while Philipp Kohlschreiber went down in straight sets to Radu Albot in the first round.
At the Cordoba Open, Londero - who was born in the city - followed up his first ATP Tour win with another victory in a remarkable contest.
Londero defeated fifth seed Nicolas Jarry on Tuesday and then, 24 hours later, saved 10 of the 10 break points he faced to see off Lorenzo Sonego 7-5 6-3.
Meanwhile, there were more shocks at the Sofia Open as top seed Karen Khachanov bowed out at the hands of Matteo Berrettini, who rallied from a set down to record an impressive 6-7 (6-8) 6-3 6-4 triumph.
Berrettini’s fellow Italian Andreas Seppi - seeded eighth for the tournament - was beaten by home hopeful Marton Fucsovics in straight sets.
Spanish duo Roberto Bautista Agut and Fernando Verdasco both progressed unscathed, though.
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